| australia | | euoniticellus sp. nlg-2014 | species | 3 | 0 |
| australia | | onitis sp. nlg-2014 | species | 3 | 0 |
| australia | | onthophagus apterus | species | 1 | 0 |
| australia | | onthophagus capella | species | 1 | 0 |
| australia | | onthophagus cuniculus | species | 1 | 0 |
| australia | | onthophagus sp. nlg-2014 | species | 2 | 0 |
| austria | | onthophagus similis | species | 1 | 0 |
| belize | | onthophagus longimanus | species | 1 | 0 |
| belize | | onthophagus nitidior | species | 1 | 0 |
| belize | | onthophagus rhinolophus | species | 1 | 0 |
| brazil | | coprophanaeus ensifer | species | 1 | 0 |
| brazil | | dichotomius schiffleri | species | 32 | 0 |
| brazil | maranhao | coprophanaeus cyanescens | species | 3 | 0 |
| bulgaria | | euoniticellus fulvus | species | 2 | 0 |
| bulgaria | | onthophagus fracticornis | species | 2 | 0 |
| bulgaria | | onthophagus ovatus | species | 1 | 0 |
| bulgaria | | onthophagus similis | species | 1 | 0 |
| bulgaria | | onthophagus taurus | species | 3 | 0 |
| cambodia | | onthophagus baolocensis | species | 1 | 0 |
| cambodia | | onthophagus cf. taurinus | species | 1 | 0 |
| cambodia | | onthophagus damaki | species | 1 | 0 |
| cambodia | | onthophagus orientalis | species | 1 | 0 |
| cambodia | | onthophagus yukae | species | 1 | 0 |
| canada | ontario | onthophagus hecate | species | 1 | 0 |
| canada | ontario | onthophagus hecate hecate | subspecies | 2 | 0 |
| canada | ontario | onthophagus orpheus | species | 3 | 0 |
| canada | ontario | onthophagus sp. bold:aah6827 | species | 1 | 0 |
| canada | ontario | onthophagus sp. bold-2016 | species | 4 | 0 |
| czech republic | usti nad labem region | onthophagus fracticornis | species | 1 | 0 |
| ecuador | | coprophanaeus sp. bmnh679884 | species | 1 | 0 |
| ecuador | | onthophagus bidentatus | species | 1 | 0 |
| ecuador | | onthophagus clypeatus | species | 1 | 0 |
| ecuador | | onthophagus haematopus | species | 1 | 0 |
| france | | onthophagus coenobita | species | 1 | 0 |
| france | | onthophagus fracticornis | species | 3 | 0 |
| france | | onthophagus lemur | species | 3 | 0 |
| france | | onthophagus maki | species | 3 | 0 |
| france | | onthophagus medius | species | 52 | 0 |
| france | | onthophagus nuchicornis | species | 3 | 0 |
| france | | onthophagus opacicollis | species | 3 | 0 |
| france | | onthophagus ruficapillus | species | 3 | 0 |
| france | | onthophagus similis | species | 4 | 0 |
| france | | onthophagus taurus | species | 3 | 0 |
| france | | onthophagus vacca | species | 39 | 0 |
| france | | onthophagus verticicornis | species | 3 | 0 |
| france | midi-pyrenees region | onthophagus vacca | species | 2 | 0 |
| france | provence-alpes-cote d'azur region | onthophagus verticicornis | species | 2 | 0 |
| france | rhone-alpes region | onthophagus verticicornis | species | 1 | 0 |
| germany | baden-wuerttemberg | onthophagus ovatus | species | 2 | 0 |
| germany | bavaria | onthophagus coenobita | species | 1 | 0 |
| germany | bavaria | onthophagus medius | species | 1 | 0 |
| germany | mecklenburg-vorpommern | onthophagus similis | species | 4 | 0 |
| germany | north rhine-westphalia | onthophagus coenobita | species | 1 | 0 |
| germany | north rhine-westphalia | onthophagus joannae | species | 1 | 0 |
| germany | north rhine-westphalia | onthophagus ovatus | species | 1 | 0 |
| germany | north rhine-westphalia | onthophagus similis | species | 8 | 0 |
| germany | north rhine-westphalia | onthophagus verticicornis | species | 2 | 0 |
| germany | saxony | onthophagus ovatus | species | 1 | 0 |
| germany | saxony-anhalt | onthophagus coenobita | species | 2 | 0 |
| germany | saxony-anhalt | onthophagus joannae | species | 2 | 0 |
| germany | saxony-anhalt | onthophagus ovatus | species | 2 | 0 |
| germany | saxony-anhalt | onthophagus similis | species | 4 | 0 |
| hong kong | | onitis falcatus | species | 1 | 0 |
| india | | onthophagus auritus | species | 1 | 0 |
| india | | onthophagus coenobita | species | 3 | 0 |
| india | | onthophagus nr. babirussa | species | 1 | 0 |
| india | | onthophagus nuchicornis | species | 4 | 0 |
| india | kerala | oniticellus cinctus | species | 2 | 0 |
| india | kerala | onthophagus dama | species | 2 | 0 |
| india | kerala | onthophagus orientalis | species | 2 | 0 |
| india | kerala | onthophagus rectecornutus | species | 2 | 0 |
| india | kerala | tiniocellus spinipes | species | 2 | 0 |
| indonesia | | onthophagus diabolicus | species | 1 | 0 |
| indonesia | | onthophagus nr. babirussa | species | 1 | 0 |
| indonesia | | onthophagus obscurior | species | 1 | 0 |
| indonesia | | onthophagus rorarius | species | 1 | 0 |
| indonesia | | onthophagus schwaneri | species | 1 | 0 |
| indonesia | | onthophagus vulpes | species | 1 | 0 |
| indonesia | | tiniocellus sarawacus | species | 1 | 0 |
| laos | | onthophagus bonasus | species | 1 | 0 |
| laos | | onthophagus cf. jeannelianus | species | 1 | 0 |
| laos | | onthophagus cf. tragus | species | 2 | 0 |
| laos | | onthophagus gracilipes | species | 1 | 0 |
| laos | | onthophagus piyawati | species | 1 | 0 |
| laos | | onthophagus tricornis | species | 1 | 0 |
| laos | | parachorius sp. 'longipennis' | species | 2 | 0 |
| laos | | parachorius sp. sp195 | species | 4 | 0 |
| laos | | parachorius thomsoni | species | 4 | 0 |
| laos | | scaptodera rhadamistus | species | 1 | 0 |
| madagascar | | digitonthophagus gazella | species | 1 | 0 |
| madagascar | | helictopleurus quadripunctatus | species | 1 | 0 |
| madagascar | | heterosyphus sicardi | species | 1 | 0 |
| malaysia | | onthophagus cervicapra | species | 1 | 0 |
| malaysia | | onthophagus crassicollis | species | 1 | 0 |
| malaysia | | onthophagus falculatus | species | 1 | 0 |
| malaysia | sabah | onthophagus cf. deflexicollis bold:ack0362 | species | 1 | 0 |
| malaysia | sabah | onthophagus nr. peninsularis bold:ack0339 | species | 2 | 0 |
| malaysia | sabah | onthophagus ochromerus | species | 1 | 0 |
| malaysia | sabah | onthophagus sp. bold:acj9390 | species | 1 | 0 |
| malaysia | sabah | onthophagus variolaris | species | 1 | 0 |
| morocco | | onthophagus andalusicus | species | 3 | 0 |
| morocco | | onthophagus nebulosus | species | 1 | 0 |
| morocco | | onthophagus vacca | species | 34 | 0 |
| myanmar | tanintharyi division | catharsius molossus | species | 1 | 0 |
| namibia | | pachysoma bennigseni | species | 4 | 0 |
| namibia | | pachysoma denticollis | species | 4 | 0 |
| namibia | | pachysoma fitzsimonsi | species | 2 | 0 |
| namibia | | pachysoma gariepinus | species | 7 | 0 |
| namibia | | pachysoma rodriguesi | species | 3 | 0 |
| namibia | | pachysoma rotundigenus | species | 3 | 0 |
| namibia | | pachysoma schinzi | species | 3 | 0 |
| namibia | | pachysoma valeflorae | species | 2 | 0 |
| nepal | | onthophagus sp. bmnh670919 | species | 2 | 0 |
| paraguay | alto paraguay department | bolbites onitoides | species | 2 | 0 |
| poland | | caccobius schreberi | species | 1 | 0 |
| romania | | euoniticellus fulvus | species | 1 | 0 |
| romania | | onthophagus fracticornis | species | 2 | 0 |
| romania | | onthophagus ruficapillus | species | 3 | 0 |
| romania | | onthophagus similis | species | 1 | 0 |
| south africa | | caccobius nigritulus | species | 1 | 0 |
| south africa | | cheironitis hoplosternus | species | 1 | 0 |
| south africa | | cleptocaccobius convexifrons | species | 2 | 0 |
| south africa | | drepanocerus kirbyi | species | 1 | 0 |
| south africa | | euoniticellus intermedius | species | 1 | 0 |
| south africa | | frankenbergerius forcipatus | species | 3 | 0 |
| south africa | | heliocopris hamadryas | species | 2 | 0 |
| south africa | | heteronitis castelnaui | species | 1 | 0 |
| south africa | | liatongus militaris | species | 1 | 0 |
| south africa | | liatongus sp. amgs 80886 | species | 1 | 0 |
| south africa | | liatongus sp. amgs 80887 | species | 1 | 0 |
| south africa | | liatongus sp. amgs 80888 | species | 1 | 0 |
| south africa | | milichus apicalis | species | 1 | 0 |
| south africa | | neateuchus proboscideus | species | 2 | 0 |
| south africa | | oniticellus egregius | species | 1 | 0 |
| south africa | | onitis alexis | species | 1 | 0 |
| south africa | | onitis fulgidus | species | 1 | 0 |
| south africa | | onthophagus alcyon | species | 1 | 0 |
| south africa | | onthophagus bicallosus | species | 1 | 0 |
| south africa | | onthophagus carbonarius | species | 2 | 0 |
| south africa | | onthophagus crassicollis | species | 2 | 0 |
| south africa | | onthophagus fimetarius | species | 1 | 0 |
| south africa | | onthophagus giraffa | species | 2 | 0 |
| south africa | | onthophagus minutus | species | 2 | 0 |
| south africa | | onthophagus pullus | species | 1 | 0 |
| south africa | | onthophagus sp. amgs 80889 | species | 1 | 0 |
| south africa | | onthophagus sp. amgs 80890 | species | 1 | 0 |
| south africa | | onthophagus sp. amgs 80891 | species | 1 | 0 |
| south africa | | onthophagus sp. amgs 80896 | species | 1 | 0 |
| south africa | | onthophagus sp. amgs 80897 | species | 1 | 0 |
| south africa | | onthophagus sp. amgs 80900 | species | 1 | 0 |
| south africa | | onthophagus sp. amgs 80926 | species | 1 | 0 |
| south africa | | onthophagus sp. amgs 80931 | species | 1 | 0 |
| south africa | | onthophagus sp. amgs 80932 | species | 1 | 0 |
| south africa | | onthophagus sp. bmnh679869 | species | 1 | 0 |
| south africa | | onthophagus vacca | species | 2 | 0 |
| south africa | | pachysoma aesculapius | species | 2 | 0 |
| south africa | | pachysoma bennigseni | species | 1 | 0 |
| south africa | | pachysoma endroeydi | species | 3 | 0 |
| south africa | | pachysoma glentoni | species | 3 | 0 |
| south africa | | pachysoma hippocrates | species | 10 | 0 |
| south africa | | pachysoma striatus | species | 3 | 0 |
| south africa | | phalops ardea | species | 1 | 0 |
| south africa | | sarophorus sp. sar01 | species | 1 | 0 |
| south africa | | sarophorus tuberculatus | species | 3 | 0 |
| south africa | | scarabaeus bohemani | species | 2 | 0 |
| south africa | | scarabaeus flavicornis | species | 2 | 0 |
| south africa | | scarabaeus rubripennis | species | 2 | 0 |
| south africa | | tiniocellus spinipes | species | 1 | 0 |
| south africa | | tragiscus dimidiatus | species | 1 | 0 |
| south africa | | xinidium sp. xin01 | species | 1 | 0 |
| south korea | | onthophagus atripennis | species | 3 | 0 |
| south korea | gyeonggi province | onthophagus sp. 01 jhy-2013 | species | 1 | 0 |
| south korea | gyeonggi province | onthophagus sp. 03 jhy-2013 | species | 1 | 0 |
| south korea | seoul city | onthophagus sp. 02 jhy-2013 | species | 1 | 0 |
| spain | | bubas bison | species | 1 | 0 |
| spain | | bubas bubalus | species | 1 | 0 |
| spain | | caccobius schreberi | species | 1 | 0 |
| spain | | euoniticellus fulvus | species | 1 | 0 |
| spain | | onthophagus similis | species | 1 | 0 |
| thailand | | onthophagus orientalis | species | 6 | 0 |
| thailand | | onthophagus vulpes | species | 3 | 0 |
| united states | georgia | phanaeus vindex | species | 1 | 0 |
| united states | hawaii | onthophagus sagittarius | species | 1 | 0 |
| united states | new york | onthophagus orpheus | species | 1 | 0 |
| united states | north carolina | onthophagus taurus | species | 31672 | 0 |