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dendroctonus frontalis infection by the dd-136 strain of neoaplectana carpocapsae and its bacterial complex.the dd-136 strain of neoaplectana carpocapsae weiser (steinernematidae) after spray application to pine bark in 0.1% formalin plus wetting agent entered pine bark beetle tunnels and killed 44% of the brood and adults of dendroctonus frontalis zimmermann at 18 and 26 c, 60% relative hummidity and at ambient temperatures and humidities.197019322322
fine structure of the prothoracic mycangium, a chamber for the culture of symbiotic fungi, in the southern pine beetle, dendroctonus frontalis.the ultrastructure of the prothoracic mycangium of female dendroctonus frontalis is examined. the mycangium consists of a cuticular invagination within which symbiotic fungi are cultured by the pine beetle and transported to new host trees. secretions from two types of gland cells pass into the mycangial lumen. the plasma membrane of type-1 cells is invaginated to form an enclosed extracellular cavity. the secretory product passes into the cavity, then through fine cylindrical channels into an e ...197118631556
improved white's solution for surface sterilization of pupae of dendroctonus frontalis. 19725085821
dimorphism in ceratocystis minor var. barrasii.ceratocystis minor var. barrasii, a fungus associated with the southern pine beetle, dendroctonus frontalis, exhibits dimorphism which is influenced by the concentration of both carbon dioxide and phosphate. the mycelium produces blastospores from the tips and sides of its hyphae when the concentration of carbon dioxide is high, and these blastospores germinate if the concentration of phosphate is above about 10(-2) mm, whereas below this concentration they bud.1977600307
redescription and life history of contortylenchus brevicomi, a parasite of the southern pine beetle dendroctonus frontalis.larval and adult life stages are described for contortylenchus brevicomi (massey) rühm parasitizing a mississippi population of dendroctonus frontalis, the southern pine beetle. fourth-stage larvae and free-living adult females of this species are identified and described for the first time. the life cycle of c. brevicomi can be reconstructed from this study. the adult female nematode lays eggs in a mature beetle. larval development progresses within the hemocoel until fourth-stage larvae exit t ...198019300698
effect of the nematode contortylenchus brevicomi on gallery construction and fecundity of the southern pine beetle.field-collected dendroctonus frontalis were reared in a controlled environment. male-female beetle pairs retrieved from galleries 1, 2, or 5 wk after introduction into pine bolts were examined for nematode parasites. data were obtained for each pair on gallery length, egg niche construction, egg viability, and progeny survival. in a separate study, beetle pairs were reared under laboratory conditions for 10 wk. the number of emerged adult progeny of each pair was recorded. contortylenchus brevic ...198019300703
niche breadth and resource partitioning by four sympatric species of bark beetles (coleoptera: scolytidae).standing loblolly pines in southeastern north america are colonized by four sympatric species of bark beetles: dendroctonus frontalis (zimm.), ips calligraphus (germ.), i. grandicollis (eichh.) and i. avulsus (eichh.). the beetles compete for the limited amount of phloem tissue used as a site for reproduction. using indices of niche breadth and niche overlap determined from the surface areas attacked, the interaction of colonizing beetle species in partitioning resources in entire trees and with ...198128309925
defensive mechanisms of loblolly and shortleaf pine against attack by southern pine beetle,dendroctonus frontalis zimmermann, and its fungal associate,ceratocystis minor (hedgecock) hunt.loblolly and shortleaf pine growing on a single site in the north carolina piedmont were examined to determine similarities and differences in their defensive mechanisms against the southern pine beetle,dendroctonus frontalis zimmermann, and its fungal associate,ceratocystis minor (hedgecock) hunt. both species responded to wounding and fungal inoculation by forming a hypersensitive lesion around the wound site. there were significantly less soluble sugars and more monoterpenes in the lesion tis ...198624307119
evidence of competition for photosynthates between growth processes and oleoresin synthesis in pinus taeda l.a group of 10 pines (pinus taeda l.) in a 35-year-old stand in central louisiana was sampled periodically from march through december 1984 for oleoresin flow from small wounds. oleoresin yield was closely associated with periods of calculated moderate soil water deficit and presumed growth reduction. the period of maximum oleoresin flow coincided with the period of calculated maximum soil water depletion. calculated potential evapotranspiration reached a maximum in late june, but water deficits ...198614975863
fungi associated with the southern pine beetle: avoidance of induced defense response in loblolly pine.loblolly pines (pinus taeda) are rapidly killed by colonizing southern pine beetle (dendroctonus frontalis). the female beetles carry two species of fungi (ceratocystis minor var. barrasii and an unnamed basidiomycete) within a mycangium. the insects are also frequently associated with a blue-staining form of c. minor. these fungi are inoculated into the tree during colonization. the tree has an induced defensive response that involves resin soaking and necrosis of affected tissue isolating the ...198728312476
antennnal olfactory and behavioral response of southern pine beetle,dendroctonus frontalis, to analogs of its aggregation pheromone frontalin.antennal olfactory (electroantennogram) and laboratory and field behavioral tests were carried out on the response ofdendroctonus frontalis to its aggregation pheromone frontalin and analogs. the analogs were compounds modified by altering the position and methyl groups and/or by their deletion. any modification to the frontalin structure significantly reduced both the antennal olfactory and behavioral response byd. frontalis. beetle response, although significantly reduced, was elicited at the ...198824276205
repellent properties of the host compound 4-allylanisole to the southern pine beetle.the phenylpropanoid 4-allylanisole is a compound produced by loblolly pines (pinus taeda l.), an abundant species in southern pine forests and a preferred host of southern pine beetle (dendroctonus frontalis zimmermann). repellency of individual beetles was demonstrated in laboratory behavioral assays ofd. frontalis and other scolytids. inhibition was demonstrated in natural populations ofd. frontalis using baited traps. in both tests, response to the inhibitory pheromone verbenone was used for ...199424242654
predation and bark beetle dynamics.bark beetle populations may undergo dramatic fluctuations and are often important pests in coniferous forests. their dynamics are thought to be primarily driven by factors affecting the resistance of the host tree to attack, i.e., bottom-up forces, while natural enemies are usually assigned a minor role in these systems. i present behavioral experiments that suggest that the clerid beetle thanasimus dubius may be an important source of mortality for the bark beetle dendroctonus frontalis during ...199728307375
dynamical role of predators in population cycles of a forest insect: an experimental testpopulation cycles occur frequently in forest insects. time-series analysis of fluctuations in one such insect, the southern pine beetle (dendroctonus frontalis), suggests that beetle dynamics are dominated by an ecological process acting in a delayed density-dependent manner. the hypothesis that delayed density dependence in this insect results from its interaction with predators was tested with a long-term predator-exclusion experiment. predator-imposed mortality was negligible during the incre ...199910446053
effects of mass inoculation on induced oleoresin response in intensively managed loblolly pine.oleoresin flow is an important factor in the resistance of pines to attack by southern pine beetle, dendroctonus frontalis zimm., and its associated fungi. abiotic factors, such as nutrient supply and water relations, have the potential to modify this plant-insect-fungus interaction; however, little is known of the effects of inoculation with beetle-associated fungi on oleoresin flow. we observed that constitutive and induced resin yield in loblolly pine, pinus taeda l., were affected by either ...200515805088
effects of tree phytochemistry on the interactions among endophloedic fungi associated with the southern pine beetle.we examined the interaction between host trees and fungi associated with a tree-killing bark beetle, dendroctonus frontalis. we evaluated (1) the response of four pinus species to fungal invasion and (2) the effects of plant secondary metabolites on primary growth of and secondary colonization of three consistent fungal associates. two of these fungi, entomocorticium sp. a and ophiostoma ranaculosum, are obligate mutualists with d. frontalis, and the third associate is a blue-staining fungus, o. ...200515898500
electrophysiological and behavioral responses of dendroctonus frontalis (coleoptera: curculionidae) to volatiles isolated from conspecifics.olfactory sensitivity of the southern pine beetle, dendroctonus frontalis zimmermann, to compounds isolated from the mid/hindguts of newly emerged conspecific adults was assayed with coupled gas chromatography-electroantennographic detection. all previously reported pheromones for d. frontalis plus eight additional compounds (fenchyl alcohol, myrtenal, cis-verbenol, trans-pinocarveol, acetophenone, trans-myrtanol, cis-myrtanol, and 2-phenylethanol) consistently elicited antennal responses from a ...200516539134
fitness consequences of pheromone production and host selection strategies in a tree-killing bark beetle (coleoptera: curculionidae: scolytinae).timing of arrival at a resource often determines an individual's reproductive success. tree-killing bark beetles can reproduce in healthy trees by attacking in adequate numbers to overcome host defences that could otherwise be lethal. this process is mediated by aggregation and antiaggregation pheromones. beetles that arrive early in such a "mass attack" must contend with undiminished tree defences, and produce enough pheromones to attract more beetles, but have a head start on gallery construct ...200616609873
antagonisms, mutualisms and commensalisms affect outbreak dynamics of the southern pine beetle.feedback from community interactions involving mutualisms are a rarely explored mechanism for generating complex population dynamics. we examined the effects of two linked mutualisms on the population dynamics of a beetle that exhibits outbreak dynamics. one mutualism involves an obligate association between the bark beetle, dendroctonus frontalis and two mycangial fungi. the second mutualism involves tarsonemus mites that are phoretic on d. frontalis ("commensal"), and a blue-staining fungus, o ...200616323015
development of a satellite-based hazard rating system for dendroctonus frontallis (coleoptera: scolytidae) in the ouachita mountains of arkansas.the southern pine beetle, dendroctonus frontalis zimmermann (coleoptera: scolytidae), is the most damaging forest insect pest of pines (pinus spp.) throughout the southeastern united states. hazard rating schemes have been developed for d. frontalis, but for these schemes to be accurate and effective, they require extensive on-site measurements of stand attributes such as host density, age, and basal area. we developed a stand hazard-rating scheme for several watersheds in the ouachita highlands ...200717461062
impact of minimum winter temperatures on the population dynamics of dendroctonus frontalis.predicting population dynamics is a fundamental problem in applied ecology. temperature is a potential driver of short-term population dynamics, and temperature data are widely available, but we generally lack validated models to predict dynamics based upon temperatures. a generalized approach involves estimating the temperatures experienced by a population, characterizing the demographic consequences of physiological responses to temperature, and testing for predicted effects on abundance. we e ...200717494404
evidence that (+)-endo-brevicomin is a male-produced component of the southern pine beetle aggregation pheromone.previous research indicated that the aggregation pheromone of the southern pine beetle, dendroctonus frontalis, is produced only by females, the sex that initiates attacks. we provide evidence indicating that secondarily arriving males augment mass aggregation by releasing the attractive synergist (+)-endo-brevicomin. healthy pines artificially infested with both sexes of d. frontalis were significantly more attractive to conspecifics than trees infested solely with females. coupled gas chromato ...200717629772
attraction of the southern pine beetle, dendroctonus frontalis, to pheromone components of the western pine beetle, dendroctonus brevicomis (coleoptera: curculionidae: scolytinae), in an allopatric zone.subtle differences in pheromone components of sympatric species should be attractive only to the producing species and unattractive or repellent to the nonproducing species, and thereby maintain reproductive isolation and reduce competition between species. bark beetles dendroctonus brevicomis and d. frontalis (coleoptera: curculionidae) are known to have common pheromone components, except for exo-brevicomin, which is produced by d. brevicomis. we predicted that d. frontalis would not respond t ...200818348798
influence of elevation on bark beetle (coleoptera: curculionidae, scolytinae) community structure and flight periodicity in ponderosa pine forests of arizona.we examined abundance and flight periodicity of five ips and six dendroctonus species (coleoptera: curculionidae, scolytinae) among three different elevation bands in ponderosa pine (pinus ponderosa douglas ex. lawson) forests of northcentral arizona. bark beetle populations were monitored at 10 sites in each of three elevation bands (low: 1,600-1,736 m; middle: 2,058-2,230 m; high: 2,505-2,651 m) for 3 yr (2004-2006) using pheromone-baited lindgren funnel traps. trap contents were collected wee ...200818348801
fine-scale genetic population structure of southern pine beetle (coleoptera: curculionidae) in mississippi forests.the southern pine beetle, dendroctonus frontalis zimmerman, is the most destructive insect pest of pine forests in the southeastern united states, mexico, and central america. southern pine beetle aggressively attacks pine trees, and when in epidemic stages, they are capable of killing even the most healthy pine trees in a short period of time. despite the amount of destruction caused by the southern pine beetle and the amount of monetary loss faced by the timber industry and recreation, the pop ...200818348820
resin flow responses to fertilization, wounding and fungal inoculation in loblolly pine (pinus taeda) in north carolina.resin flow is the primary means of natural defense against southern pine beetle (dendroctonus frontalis zimm.), the most important insect pest of pinus spp. in the southern united states. as a result, factors affecting resin flow are of interest to researchers and forest managers. we examined the influence of fertilization, artificial wounding and fungal inoculation on resin flow in 6- and 12-year-old stands of loblolly pine (pinus taeda l.) and determined the extent of that influence within and ...200818381265
global and comparative protein profiles of the pronotum of the southern pine beetle, dendroctonus frontalis.the southern pine beetle (dendroctonus frontalis zimmermann) kills all pines within its range and is among the most important forest pest species in the us. using a specialized mycangium surrounded by gland cells in the pronotum, adult females culture, transport, and inoculate two fungi into beetle galleries during oviposition. these fungal symbionts, to varying degrees, exclude antagonistic fungi and provide nutrients to larvae. however, the mechanisms (e.g. secreted antibiotic chemicals or nut ...200818477241
temperature extremes, density dependence, and southern pine beetle (coleoptera: curculionidae) population dynamics in east texas.previous studies of the southern pine beetle, dendroctonus frontalis zimm., established that its population in east texas responds to a delayed density-dependent process, whereas no clear role of climate has been determined. we tested two biological hypotheses for the influence of extreme temperatures on annual southern pine beetle population growth in the context of four alternative hypotheses for density-dependent population regulation. the significance of climate variables and their interacti ...200818559170
field response of dendroctonus frontalis (coleoptera: scolytinae) to synthetic semiochemicals in chiapas, mexico.dendroctonus frontalis zimmermann (coleoptera: scolytinae) is the most serious pest of pines (pinus spp.) in mexico. conspecifics are attracted to trees undergoing colonization by the aggregation pheromone frontalin, which is synergized by odors of pine oleoresin released from beetle-damaged host tissue. synthetic racemic frontalin combined with turpentine has been the operational bait used in traps for monitoring populations of d. frontalis in mexico as well as the united states. recently, race ...200819133462
efficacy of two systemic insecticides injected into loblolly pine for protection against southern pine bark beetles (coleoptera: curculionidae).we evaluated the efficacy of systemic insecticides emamectin benzoate and fipronil for preventing mortality of individual loblolly pines, pinus taeda l., as a result of attacks by southern pine bark beetles (coleoptera: curculionidae, scolytinae) for two consecutive years in mississippi (2005-2006) and alabama (2006-2007). trees were injected once in the spring of 2005 (mississippi) or 2006 (alabama) and then were baited with species-specific bark beetle lures several weeks later. the southern p ...200919610420
efficacy of verbenone for protecting ponderosa pine stands from western pine beetle (coleoptera: curculionidae: scolytinae) attack in california.the western pine beetle, dendroctonus brevicomis leconte (coleoptera: curculionidae: scolytinae), is a major cause of ponderosa pine, pinus ponderosa dougl. ex laws., mortality in much of western north america. currently, techniques for managing d. brevicomis infestations are limited. verbenone (4,6,6-trimethylbicyclo [3.1.1] hept-3-en-2-one) is an antiaggregation pheromone of several dendroctonus spp., including d. brevicomis, and it has been registered as a biopesticide for control of mountain ...200919886449
spatial displacement of release point can enhance activity of an attractant pheromone synergist of a bark beetle.flight responses of the southern pine beetle, dendroctonus frontalis zimmermann, to widely-spaced (>130 m) traps baited with pine volatiles (in turpentine) and the female-produced pheromone component frontalin were enhanced when a bait containing the male pheromone component (+)-endo-brevicomin was attached directly to the trap. however, displacing this bait 4-16 m horizontally from the trap significantly increased its synergistic effect. (+)-endo-brevicomin enhanced catch to the same degree whe ...200919902305
mycangimycin, a polyene peroxide from a mutualist streptomyces sp.a mutualist actinomycete of the southern pine beetle, dendroctonus frontalis, produces a polyene peroxide with pronounced antifungal activity. its structure, absolute configuration, and biological activity were determined by spectral analysis, chemical modification followed by the modified mosher method, and growth inhibitory assays, respectively.200919125624
modeling the impacts of two bark beetle species under a warming climate in the southwestern usa: ecological and economic consequences.predicted climate warming is expected to have profound effects on bark beetle population dynamics in the southwestern united states. temperature-mediated effects may include increases in developmental rates, generations per year, and changes in habitat suitability. as a result, the impacts of dendroctonus frontalis and dendroctonus mexicanus on forest resources are likely subject to amplification. to assess the implications of such change, we evaluated the generations per year of these species u ...200919680717
a conceptual framework: redefining forest soil's critical acid loads under a changing climate.federal agencies of several nations have or are currently developing guidelines for critical forest soil acid loads. these guidelines are used to establish regulations designed to maintain atmospheric acid inputs below levels shown to damage forests and streams. traditionally, when the critical soil acid load exceeds the amount of acid that the ecosystem can absorb, it is believed to potentially impair forest health. the excess over the critical soil acid load is termed the exceedance, and the l ...201020045233
description of parasitorhabditis frontali n. sp. (nemata: rhabditida) from dendroctonus frontalis zimmermann (coleoptera: scolytidae).a new parasitorhabditis species with males and females was discovered from the southern pine beetle dendroctonus frontalis and its galleries in loblolly pine, pinus taeda, growing in mississippi. females of the new species have a cupola-shaped tail with a small spike; males possess a 2 + (3+2) + 3 ray pattern on the tail fan with ray 10 reaching the margin, and a distinctive stomatal tooth. parasitorhabditis frontali n. sp. has some similarities to p. hylurgi massey, 1974 from hylurgops pinifex ...201022736836
presence and diversity of streptomyces in dendroctonus and sympatric bark beetle galleries across north america.recent studies have revealed several examples of intimate associations between insects and actinobacteria, including the southern pine beetle dendroctonus frontalis and the spruce beetle dendroctonus rufipennis. here, we surveyed streptomyces actinobacteria co-occurring with 10 species of dendroctonus bark beetles across the united states, using both phylogenetic and community ecology approaches. from these 10 species, and 19 other scolytine beetles that occur in the same trees, we obtained 154 ...201121249352
variable responses by southern pine beetle, dendroctonus frontalis zimmermann, to the pheromone component endo-brevicomin: influence of enantiomeric composition, release rate, and proximity to infestations.the male-produced bicyclic acetal endo-brevicomin is a component of the pheromone blend that mediates colonization of host pines by the bark beetle dendroctonus frontalis zimmermann. efforts to identify its behavioral function have been complicated by contrasting reports that it either enhances or reduces attraction of flying beetles. our studies failed to support the hypothesis that this published variability is due to differences in release rate and/or the enantiomeric composition [i.e., the b ...201121465266
temperature alters the relative abundance and population growth rates of species within the dendroctonus frontalis (coleoptera: curculionidae) community.temperature has strong effects on metabolic processes of individuals and demographics of populations, but effects on ecological communities are not well known. many economically and ecologically important pest species have obligate associations with other organisms; therefore, effects of temperature on these species might be mediated by strong interactions. the southern pine beetle (dendroctonus frontalis zimmermann) harbors a rich community of phoretic mites and fungi that are linked by many s ...201122251683
genetic heterogeneity in a cyclical forest pest, the southern pine beetle, dendroctonus frontalis, is differentiated into east and west groups in the southeastern united states.the southern pine beetle, dendroctonus frontalis zimmerman (coleoptera: curculionidae) is an economically important pest species throughout the southeastern united states, arizona, mexico, and central america. previous research identified population structure among widely distant locations, yet failed to detect population structure among national forests in the state of mississippi. this study uses microsatellite variation throughout the southeastern united states to compare the southern pine be ...201122220595
molecular evidence of facultative intraguild predation by monochamus titillator larvae (coleoptera: cerambycidae) on members of the southern pine beetle guild.the southern pine bark beetle guild (spbg) is arguably the most destructive group of forest insects in the southeastern usa. this guild contains five species of bark beetles (coleoptera: curculionidae: scolytinae): dendroctonus frontalis, dendroctonus terebrans, ips avulsus, ips calligraphus, and ips grandicollis. a diverse community of illicit receivers is attracted to pheromones emitted by the spbg, including the woodborers monochamus carolinensis and monochamus titillator (coleoptera: ceramby ...201223052822
mycangia of ambrosia beetles host communities of bacteria.the research field of animal and plant symbioses is advancing from studying interactions between two species to whole communities of associates. high-throughput sequencing of microbial communities supports multiplexed sampling for statistically robust tests of hypotheses about symbiotic associations. we focus on ambrosia beetles, the increasingly damaging insects primarily associated with fungal symbionts, which have also been reported to support bacteria. to analyze the diversity, composition, ...201222546962
semiochemical emission from individual galleries of the southern pine beetle, (coleoptera: curculionidae: scolytinae), attacking standing trees.we collected, identified, and quantified volatiles arising from individual gallery entrances of the monogamous bark beetle dendroctonus frontalis zimmermann. samples were collected while the insects were mass attacking mature loblolly pines (pinus taeda l.) in an established infestation in western mississippi, 1 august through 3 october 2005. following volatile sample collection, the entrances were dissected and categorized according to those that 1) contained a solitary female (the gallery init ...201222420266
southern pine beetle, dendroctonus frontalis, antennal and behavioral responses to nonhost leaf and bark volatiles.a growing body of evidence suggests that bark beetles detect and avoid release points of volatile compounds associated with nonhost species, and thus such nonhost volatiles may have potential utility in the management of bark beetles. we used a coupled gas chromatograph-electroantennographic detector (gc-ead) to assay the olfactory sensitivity of the southern pine beetle, dendroctonus frontalis zimmermann, to volatiles from leaves and bark of eight species of nonhost angiosperm trees that are co ...201323460417
incorporating carbon storage into the optimal management of forest insect pests: a case study of the southern pine beetle (dendroctonus frontalis zimmerman) in the new jersey pinelands.forest insect pest disturbance is increasing in certain areas of north america as many insect species, such as the southern pine beetle, expand their range due to a warming climate. because insect pests are beginning to occupy forests that are managed for multiple uses and have not been managed for pests before, it is becoming increasingly important to determine how forests should be managed for pests when non-timber ecosystem services are considered in addition to traditional costs and revenues ...201424938795
use of acoustics to deter bark beetles from entering tree material.acoustic technology is a potential tool to protect wood materials and eventually live trees from colonization by bark beetles. bark beetles such as the southern pine beetle dendroctonus frontalis, western pine beetle d. brevicomis and pine engraver ips pini (coleoptera: curculionidae) use chemical and acoustic cues to communicate and to locate potential mates and host trees. in this study, the efficacy of sound treatments on d. frontalis, d. brevicomis and i. pini entry into tree materials was t ...201424376044
pheromone-mediated mate location and discrimination by two syntopic sibling species of dendroctonus bark beetles in chiapas, mexico.where their geographic and host ranges overlap, sibling species of tree-killing bark beetles may simultaneously attack and reproduce on the same hosts. however, sustainability of these potentially mutually beneficial associations demands effective prezygotic reproductive isolation mechanisms between the interacting species. the pine bark beetle, dendroctonus frontalis zimmermann, is syntopic in the central american region with a recently described sibling species, dendroctonus mesoamericanus arm ...201526255157
western pine beetle populations in arizona and california differ in the composition of their aggregation pheromones.we compared pheromone production and response for populations of western pine beetle, dendroctonus brevicomis leconte, from sites in northern arizona and northern california. volatiles were collected from individuals of both sexes that had mined as a pair in a pinus ponderosa log for 1 d, and they were subsequently analyzed by gas chromatography coupled to mass-spectrometry. principal component analysis of quantities of dendroctonus pheromone components indicated strong site-associated clusterin ...201627125814
intra-annual variation in responses by flying southern pine beetles (coleoptera: curculionidae: scolytinae) to pheromone component endo-brevicomin.the southern pine beetle dendroctonus frontalis zimmermann (coleoptera: curculionidae: scolytinae) is attracted to an aggregation pheromone that includes the multifunctional pheromone component endo-brevicomin. the effect of endo-brevicomin on attractive lures varies from strong enhancement to reduction of beetle attraction depending upon release rate, lure component spacing, and proximity of beetle infestations. anecdotal observations have further suggested that the effects of endo-brevicomin v ...201627207264
responses by dendroctonus frontalis and dendroctonus mesoamericanus (coleoptera: curculionidae) to semiochemical lures in chiapas, mexico: possible roles of pheromones during joint host attacks.in southern mexico and central america, the southern pine beetle dendroctonus frontalis zimmermann (coleoptera: curculionidae: scolytinae) commonly colonizes host trees simultaneously with dendroctonus mesoamericanus armendáriz-toledano and sullivan, a recently described sibling species. we hypothesized that cross-species pheromone responses by host-seeking beetles might mediate joint mass attack, bole partitioning, and reproductive isolation between the species. previous studies had indicated t ...201626803816
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