Path
rootcellular organisms
eukaryota
opisthokonta
metazoa
eumetazoa
bilateria
coelomata
protostomia
panarthropoda
arthropoda
mandibulata
pancrustacea
hexapoda
insecta
dicondylia
pterygota
neoptera
endopterygota
coleoptera
adephaga
caraboidea
carabidae
trechinae
bembidiini
bembidion
princidium subgenus complex
actedium
bembidion pallidipenne