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a new species of phrynopus from the northeastern andes of peru, its phylogenetic position, and notes on the relationships of holoadeninae (anura: craugastoridae).we report the discovery of a geographically disjunct and morphologically distinctive species of direct-developing frog of the genus phrynopus (phrynopus mariellaleo sp. nov.) that changes considerably our understanding of the distribution of species in this andean genus. the type locality lies on a subcordillera (cerro de campanario area) of the extreme northeastern portion of the cordillera central of peru, on the headwaters of the mayo river, amazonas department, at 2575 m asl (6°6'42.9''s, 77 ...201830313872
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